Geniostoma pumilum

(Hillebr.) Byng & Christenh. (2018)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Gentianales Family: Loganiaceae Genus: Geniostoma

kāmakahala [kamakahala]

Description

Key Characters:

Growth Form: Sparingly branched shrubs.

Stems: Stems sharply angled or slightly winged, glabrous or with a few scattered hairs.

Roots:

Leaves: Leaves simple. Opposite. Blades elliptic–obovate, 4–10.5 cm long, 2–4.5 cm wide. Apex abruptly short-acuminate. Base cuneate. Surfaces glabrous, Leaves pale and glaucous on lower surface, coriaceous at least when dry. Petioles 0–1 cm long. Stipules interpetiolar, may also be intrapetiolar, membranous, connate into a cup 5–9 mm long, distinct from petioles, caducous.

Flowers: Flowers in inflorescences terminal, flowers up to 15 in sessile cymes, peduncles 0–6 mm long, pedicels 8–11 mm long, elongating to 10–13 mm long in fruit, bracts linear–filiform, 5–7 mm long, bracteoles filiform, ca. 4 mm long, glabrous. Flowers bisexual (perfect) or functionally unisexual, actinomorphic. Calyx 5-merous, lobes subequal, sepals united at base, oblong to oblong–lanceolate, 6–11 mm long, 2–3 mm wide, imbricate, conspicuously 5–7–nerved, inner surface of calyx often with colleters at the base. Corolla 5-merous, orange, 15–16 mm long, the tube not or slightly flaring, 9–10 mm long, short–hirsute externally, the lobes narrowly deltate, 5–7 mm long, apex acute; lobes imbricate or contorted in bud. Stamens 5, alternipetalous, attached to mouth of corolla, exserted, usually recurved after anthesis, glabrous or hairy; filaments mostly short; anthers 2-celled, introrse, connective often extended to form an apical appendage, opening by longitudinal slits. Ovary superior or partly inferior, 2– or 3-locular, sometimes partitions imperfect in upper part of ovary, glabrous or slightly pubescent near apex, pistillode glabrous; ovules numerous, amphitropous, placentation axiles; style terminal, mostly short; stigma clavate, ellipsoid, to globular, sometimes ± oblong, about as large as ovary.

Fruit: Capsules ovoid–trigonous; 9–14 mm long; 6–8 mm in diameter; 3–valved; septicidal to sub–septifragal; valves obscurely to conspicuously keeled near apex; keel 0.5–1.5 mm wide; apex with a beak 1.5–2.5 mm long; maturing to green; with or without a black–purple tinge; usually becoming brown–black when over–mature. Seeds ± winged at both ends or all around; embedded in juicy orange to red pulp; numerous; ellipsoid to subglobular; intruded on the hilar side; surface mostly minutely papillose to areolate; brown to black; endosperm thick; fleshy; oily; proteinaceous; with saccharose or hemicellulose; embryo small; straight.

Ploidy:

Habitat: Occurring on hummocks in bogs and probably bog margins.

Elevation Range: 1,060–1,390 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Global Fl. 4: 145 (2018)

Other References

Wagner et al. 1990:860 (K)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Eastern wind-swept ridges, montane shrubs with forest patches on flanks. PRESERVED_SPECIMEN K. R. Wood 12287 Kauai PTBG 3/6/2007
2 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN a. s. hitchcock 15445 Kauai US 10/22/1916
3 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Montane bog PRESERVED_SPECIMEN n. tangalin, n. barca & k. kagimoto 2492 Kauai US 2/3/2011
4 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Eastern wind-swept ridges, montane shrubs with forest patches on flanks. PRESERVED_SPECIMEN K. R. Wood 12291 Kauai PTBG 3/6/2007
5 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Montane bogs w/ mixed sedges and grasses, bogs dissected w/ riparian veg of Metrosideros-Cheirodendron forest. PRESERVED_SPECIMEN K. R. Wood 11663 Kauai PTBG 12/30/2005
6 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN k. r. wood, s. p. perlman & r. gillespie 1860 Oahu US 5/2/1992
7 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron montane wet forest, bogs and riparian habitats PRESERVED_SPECIMEN K. R. Wood 17913 Kauai PTBG 8/14/2018
8 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN k. r. wood 3898 Kauai US 12/28/1994
9 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN k. r. wood, s. p. perlman & r. gillespie 1862 Kauai US 5/2/1992
10 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN k. r. wood, m. dumaran & c. thomson 4892 Kauai US 1/2/1996
11 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Royen, P. van 11722 Kauai US 9/3/1977
12 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Montane bog PRESERVED_SPECIMEN N. Tangalin, N. Barca & K. Kagimoto 2492 Kauai US 2/3/2011
13 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Perlman, S. P.; Wood, K.; Mehrhoff, L. 14603 Kauai US 12/29/1994
14 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Montane bog PRESERVED_SPECIMEN Tangalin, N.; Barca, N.; Kagimoto, K. 2492 Kauai US 2/3/2011
15 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN a. s. hitchcock 15498 Kauai US 10/22/1916
16 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wood, K. R.; Perlman, S. P.; Gillespie, R. 1860 Kauai US 5/2/1992
17 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wood, K. R.; Dumaran, M.; Thomson, C. 4892 Kauai US 1/2/1996
18 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wood, K. R. 3898 Kauai US 12/28/1994
19 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN p. van royen 11722 Oahu US 9/3/1977
20 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Perlman, S. P.; Buck, M.; Gon, S. 9206 Kauai US 5/19/1988
21 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wood, K. R.; Perlman, S. P.; Gillespie, R. 1862 Kauai US 5/2/1992
22 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN s. p. perlman, k. wood & l. mehrhoff 14603 Kauai US 12/29/1994
23 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Hitchcock, A. S. 15445 Kauai US 1916-00-00
24 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron montane wet mixed communities. PRESERVED_SPECIMEN K. R. Wood 1862 Kauai PTBG 5/2/1992
25 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN s. p. perlman, m. buck & s. gon 9206 Kauai US 5/19/1988
26 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information In open boggy areas. Metrosideros bogss and Metrosideros-Cheirodendron montane wet forest with Cibotium, Vaccinium, Myrsine, Rhynchospora, Viola, Melicope, Styphelia, Stenogyne, Drosera, Lagenifera, Dubautia paleata, Labordia, Coprosma. PRESERVED_SPECIMEN Perlman, S. 9206 Kauai BISH 5/19/1988
27 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 8884 Kauai BISH 10/20/1911
28 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 905.K Kauai BISH 7/3/1917
29 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Open Metrosideros, Myrsine and Acacia forest on soil substrate. Understory consisting of Athyrium, Dryopteris, Metrosideros, Myoporum, Myrsine, Pelea, Pipturus, Rubus and Vaccinium. PRESERVED_SPECIMEN wood, k.r. collector number: 12291 Hawaii BISH 3/6/2007
30 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information E wind-swept ridges, montane shrubs with forest patches on flanks. 100% native with Dicranopteris & Diplopterygium 1 - 2 m tall, Melicope feddei, Cheirodendron dominii, Dubautia knudsenii, D laxa, Labordia waialealae, L. pumila, Lobelia kauaiensis. PRESERVED_SPECIMEN Wood, K.R. 12291 Kauai BISH 3/6/2007
31 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 4874 Kauai BISH 3/23/1905
32 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros mixed bogs along rolling hills of pali in wind swept area w/ Carex sp, Rhynchospora, Vaccinium, Lagenifera, Viola kauaiense, Dubautia paleata. PRESERVED_SPECIMEN Wood, K.R. 3671 Kauai BISH 10/25/1994
33 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 8884 Kauai BISH 10/20/1911
34 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron montane wet forest, bogs and riparian habitats w/ Broussaisia arguta, Dubautia paleata, Labordia waialealae, L. Pumila, Polyscias waialealae, Vaccinum calycinum, Pipturus ruber, Deschampsia nubigena, Machaerina angustifolia, Melicope clusiifolia, PRESERVED_SPECIMEN Wood, K.R. 17919 Kauai BISH 8/14/2018
35 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information montane bogs; Rhynchospora chinensis, R. rugosa, Dichanthelium, Deschampsia, Machaerina, Plantago pachyphylla, Viola, Metrosideros, Dubautia waialealae, D. imbricata, D. laxa, Cheirodendron dominii, Leptecophylla tameiameiae PRESERVED_SPECIMEN Wood, K.R. 11663 Kauai BISH 12/30/2005
36 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information E wind-swept ridges, montane shrubs with forest patches on flanks. 100% native with Dicranopteris & Diplopterygium 1 - 2 m tall, Melicope feddei, Cheirodendron dominii, Dubautia knudsenii, D laxa, Labordia waialealae, L. pumila, Lobelia kauaiensis. PRESERVED_SPECIMEN Wood, K.R. 12287 Kauai BISH 3/6/2007
37 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 905.k Kauai BISH 7/3/1917
38 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information turfy ledges PRESERVED_SPECIMEN lydgate, j.m. collector number: s.n. Maui BISH
39 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Near rain swept summit. PRESERVED_SPECIMEN wood, k.r. nishek, r. collector number: 11663 Oahu BISH 12/30/2005
40 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 4874 Kauai BISH
41 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN perlman, s.p. buck, m.; gon, s. collector number: 9206 Kauai BISH 5/19/1988
42 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Bushy hummocks in bog. PRESERVED_SPECIMEN St. John, H. 23059 Kauai BISH 12/25/1947
43 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information In swamp forest PRESERVED_SPECIMEN van Royen, P. 11722 Kauai BISH 9/3/1977
44 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Bushy hummocks in bog. PRESERVED_SPECIMEN st.john, h. cowan, r.s.; webster, g.l.; wilbur, r.l. collector number: 23059 Kauai BISH 12/25/1947
45 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 5813 Kauai BISH 9/23/1909
46 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 1153.K Kauai BISH 7/3/1917
47 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information In swamp forest. PRESERVED_SPECIMEN van royen, p. davis, j.; perlman, s. collector number: 11722 Kauai BISH 9/3/1977
48 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 1153.k Kauai BISH 7/3/1917
49 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information In swamp forest. PRESERVED_SPECIMEN van Royen, P. 11722 Kauai BISH 9/3/1977
50 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros mixed bogs along rolling hills of pali in wind swept area w/ Carex sp, Rhynchospora, Vaccinium, Lagenifera, Viola kauaiense, Dubautia paleata. PRESERVED_SPECIMEN wood, k.r. perlman, s. collector number: 3671 Kauai BISH 10/25/1994
51 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 5813 Kauai BISH 9/23/1909
52 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information along stream banks. Dubautia-Sadleria shrubland/fernland, high wind summit slopes w/ Psychotria mariniana, Kadua affinis, Melicope clusiifolia, M. waialealae, Vaccinum calycinum, Coprosma kauaense, Dubautia laxa, D. paleata, D. imbricata acronaea, Broussaisia arguta, Bidens forbes PRESERVED_SPECIMEN Wood, K.R. 15470 Kauai BISH 4/3/2013
53 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 938.k Kauai BISH 7/3/1917
54 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 938.K Kauai BISH 7/3/1917
55 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Montane bogs dissected w/ riparian veg of Metrosideros-Cheirodendron forest. PRESERVED_SPECIMEN K. R. Wood 12304 Kauai PTBG 3/7/2007
56 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information In Metrosideros woodland. PRESERVED_SPECIMEN John J. Fay 218 Kauai PTBG 6/13/1974
57 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros bogs and Metrosideros-Cheirodendron montane wet forest. PRESERVED_SPECIMEN Steve Perlman 9206 Kauai PTBG 5/19/1988
58 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information sloping bogs with adjacent Metrosideros-Cheirodendron montane wet forest with small side streams and wind swept slopes PRESERVED_SPECIMEN K. R. Wood 16006 Kauai PTBG 7/30/2014
59 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron Montane Wet Forest. PRESERVED_SPECIMEN K. R. Wood 4892 Kauai PTBG 1/2/1996
60 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Dubautia-Sadleria shrubland/fernland, high wind summit slopes PRESERVED_SPECIMEN K. R. Wood 15470 Kauai PTBG 4/3/2013
61 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros mixed bogs along rolling hills of pali in wind swept area. PRESERVED_SPECIMEN K. R. Wood 3671 Kauai PTBG 10/25/1994
62 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Scattered bogs in montane wet Metrosideros-Cheirodendron forest w/ stream systems dissecting; PRESERVED_SPECIMEN K. R. Wood 3898 Kauai PTBG 12/28/1994
63 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 8273 Kauai PTBG 2/29/2000
64 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Dubautia-Sadleria shrubland-fernland PRESERVED_SPECIMEN K. R. Wood 16197 Kauai PTBG 1/7/2015
65 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 8208 Kauai PTBG 2/1/2000
66 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros mixed bogs. PRESERVED_SPECIMEN K. R. Wood 3670 Kauai PTBG 10/25/1994
67 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Montane bogs w/ mixed sedges and grasses, bogs dissected w/ riparian veg of Metrosideros-Cheirodendron forest. PRESERVED_SPECIMEN K. R. Wood 11704 Kauai PTBG 1/1/2006
68 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron shrubland. PRESERVED_SPECIMEN Steve Perlman 14603 Kauai PTBG 12/29/1994
69 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron montane wet mixed communities. PRESERVED_SPECIMEN K. R. Wood 1860 Kauai PTBG 5/2/1992
70 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Low, wet elfin cloud forest. PRESERVED_SPECIMEN David H. Lorence 6025 Kauai PTBG 6/17/1988
71 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Montane Metrosideros-Oreobolus Bog PRESERVED_SPECIMEN Natalia Tangalin 2492 Kauai PTBG 2/3/2011
72 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Scattered bogs in montane wet Metrosideros-Cheirodendron forest w/ stream systems dissecting. PRESERVED_SPECIMEN K. R. Wood 3886 Kauai PTBG 12/28/1994
73 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information scattered bogs in montane wet Metrosideros-Cheirodendron forest w/ stream systems dissecting. PRESERVED_SPECIMEN K. R. Wood 3893 Kauai PTBG 12/28/1994
74 Geniostoma pumilum (Hillebr.) Byng & Christenh. Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron montane wet forest, bogs and riparian habitats PRESERVED_SPECIMEN K. R. Wood 17919 Kauai PTBG 8/14/2018